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Cooliris is a visually appealing image browsing addon with a unique mouse interface.
Locationbar² - it's not very stylish, but it is very usable.
Relevant screenshots:
a) cleaner URLs
b) each part acts as a separate link, my mouse is over 'questions'
c) available options
Firefox 3.7 Theme
This is based on Mozilla's design mockups for Firefox 3.7.
Installing this theme isn't very simple. You'll need to first install the All-Glass Firefox extension for the Aero effects, the Personal Menu extension to hide the menubar and add the Tools button, the Stylish extension for a tweak that fixes the text, and then drag the mockup theme's *.jar file into the add-ons window to install it.
For detailed instructions: http://lifehacker.com/5322073/firefox-37-theme-makes-your-browser-look-awesome
For the Firefox 3.7 theme: http://boneyardbrew.deviantart.com/art/Firefox-3-7-Mockup-Redux-2-0-130502568
After thinking a little, the best look and feel can only be achieved with extensions that let you tweak them (i.e., everyone has preferences.) So my pick goes for Stylish.
It lets you apply CSS to almost anything that can be represented in Firefox, even XUL!
An example of it, I've messed with my history menu to make it unique, here is how it looks after some work (CSS3 on the XUL omg omg):
Okay, it's not a full-blown application. but Stylish and userstyles.org are for CSS what Greasemonkey and userscripts.org are for JavaScript, so possibilities for a "rich look and feel" are almost unlimited.
