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what are the ports is required to do the mailbox migration in On-premises to O365? what are the ports are required to do the mailbox migration in O365 -O365?

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That may depend on the type of the On-premises installation.

In general, these ports are required for Office 365 according to your needs :

  • TCP 443 - ADFS, Microsoft Online Portal (Website), Outlook 2010, Outlook 2007, Outlook Web Access (Website), SharePoint Online (Website), Lync 2010 client, CRLs (Certificate Revocation Lists) to verify SSL
  • TCP 25 - SMTP Mail routing
  • TCP 587 - SMTP relay
  • TCP 143/993 - IMAP migration tool
  • TCP 995 - POP3
  • TCP 80 and 443 - Online services directory, simple exchange migration tool, simple IMAP migration
  • STUN/UDP 3478 - Lync Online (outbound audio and video session)
  • RTC/UDP 50000-59999 - Lync Online (outbound audio and video session)
  • TCP 50040-50059 - App sharing and file transfer
  • STUN/TCP 443 - Lync Online (outbound audio, video, and app sharing session)
  • TCP 443 - Lync (Data sharing sessions)
  • TCP 5223 - Lync Mobile Push iOS only
  • TCP/UDP 5671 - when using Azure Service Bus
  • TCP 6001, 6002, 6004 - when migrating from Exchange 2003

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